In soft species of Triticum spelta L., T. compactum Host. and T.sphaerococcum Persiv. the author studied manifestation of complex quantitative determinants on basis of phenome—genome approach, when the genome-phenome interrelation are considered similarly to gene— phene. It was shown, that at identical genome composition (AABBDD) in T. aestivum L.,T.spelta L., T. compactum Host. and T. sphaerococcum Persiv. , in three last-named cultivars the quantitative determinants are controlled on tetraploid twophenome level (diploid genome—phenome), that the author suggests to reflect in genomic formulae and take into account in selection and practical work.